The death of Alexei Navalny has been confirmed by his spokesperson, but it remains unclear where the body of the Vladimir Putin critic is.

Spokesperson Kira Yarmysh said in a post on X the most prominent face of the Russian opposition to Mr Putin was “murdered” at a remote Arctic penal colony.

She said Mr Navalny’s mother, Lyudmila Navalnaya, was informed by authorities that her son had died on 16 February at 2.17pm, local time.

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Prominent Navalny ally Ivan Zhdanov added that prison officials told Mr Navalny’s mother that he had died due to “sudden death syndrome”.

But the body of the 47-year-old has not yet been located or released by authorities.

Mr Navalny’s mother was told by a prison official that her son’s body was taken to the nearby city of Salekhard as part of a probe into his death, Ms Yarmysh said.

But when they arrived at the morgue, it was closed, and workers said the body was not there.

Hours later, Ms Yarmysh said lawyers for the politician were told Mr Navalny’s body would not be handed over to his relatives until an investigation into his death had been completed.

She accused the Investigation Committee in Salekhard of “driving us around in circles and covering their tracks” as only hours before they were told the investigation had already been concluded, and nothing criminal had been established.

Russia’s Federal Penitentiary Service reported on Friday that Mr Navalny felt sick after a walk and became unconscious at the penal colony in the town of Kharp, in the Yamalo-Nenets region, which is within the Arctic Circle.

More than 340 detained in Russia

Meanwhile, more than 340 people have been arrested in Russia since the moment Mr Navalny’s death became public, according to independent human rights organisation, OVD-Info.

This included 230 people across multiple cities who were detained on Saturday.

OVD-Info said others had been detained the day before when they came to lay flowers in memory of Mr Navalny.

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Among the arrested included a priest who went to a memorial in St Petersburg to conduct a service in the politician’s memory.

In Moscow, social media footage showed a large group of people chanting “shame” as police dragged a screaming woman from the crowd.

Putin ‘should be held accountable’

Hours after Mr Navalny’s death was reported, his wife, Yulia Navalnaya, made a dramatic appearance at a security conference in Germany where many world leaders had gathered.

She said that she was unsure if she could believe the news from official Russian sources, “but if this is true, I want Putin and everyone around Putin, Putin’s friends, his government to know that they will bear responsibility for what they did to our country, to my family and to my husband”.

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Reacting, Labour leader Sir Keir Starmer said he was “moved” by Mrs Navalnaya’s words, adding that Russia has to be held to account.

It came after the UK’s Foreign Secretary Lord Cameron said: “We should hold Putin accountable for this. And no one should be in any doubt about the dreadful nature of Putin’s regime in Russia after what has just happened.”

Foreign ministers of the G7 – made up of Canada, France, Germany, Italy, Japan, the UK and US – have called on Russia to fully clarify the circumstances of Mr Navalny’s death.

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